Acis (plant)
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''Acis '' is a genus of
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
,
herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition of t ...
and bulbous plants in the amaryllis family (
Amaryllidaceae The Amaryllidaceae are a family of herbaceous, mainly perennial and bulbous (rarely rhizomatous) flowering plants in the monocot order Asparagales. The family takes its name from the genus ''Amaryllis'' and is commonly known as the amaryllis fa ...
, subfamily
Amaryllidoideae Amaryllidoideae (Amaryllidaceae ''s.s.'', amaryllids) is a subfamily of monocot flowering plants in the family Amaryllidaceae, order Asparagales. The most recent APG classification, APG III, takes a broad view of the Amaryllidaceae, which then ...
). The genus consists of nine species distributed in Europe and Northern Africa. ''Acis'' was previously included in '' Leucojum''; both genera are known as snowflakes.


Description

''Acis'' species are
perennial A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also wide ...
bulb In botany, a bulb is structurally a short stem with fleshy leaves or leaf basesBell, A.D. 1997. ''Plant form: an illustrated guide to flowering plant morphology''. Oxford University Press, Oxford, U.K. that function as food storage organs duri ...
ous plants. The flowers have six equally sized
tepal A tepal is one of the outer parts of a flower (collectively the perianth). The term is used when these parts cannot easily be classified as either sepals or petals. This may be because the parts of the perianth are undifferentiated (i.e. of very ...
s, unlike the related genus ''
Galanthus ''Galanthus'' (from Ancient Greek , (, "milk") + (, "flower")), or snowdrop, is a small genus of approximately 20 species of bulbous perennial herbaceous plants in the family Amaryllidaceae. The plants have two linear leaves and a single ...
'' (snowdrops) in which the inner three tepals are shorter than the outer three. The tepals are unmarked, differing in this respect from '' Leucojum''. Most species have white tepals, although those of '' Acis rosea'' are pale pink. ''Acis'' species are relatively short, up to in '' Acis autumnalis'' and in ''
Acis tingitana ''Acis tingitana'' is a species of flowering plant in the family Amaryllidaceae, native to north Morocco. Rare in cultivation, it is not fully frost-hardy. Description ''Acis tingitana'' is a bulbous flowering plant. Forms described under the ...
''. The leaves are narrow; very narrow (filiform) in '' Acis trichophylla''. The flower stalks (scapes) are solid.


Taxonomy

''Acis'' was first differentiated from the genus '' Leucojum'' by
Richard Anthony Salisbury Richard Anthony Salisbury, FRS (born Richard Anthony Markham; 2 May 1761 – 23 March 1829) was a British botanist. While he carried out valuable work in horticultural and botanical sciences, several bitter disputes caused him to be ostracised ...
in ''
The Paradisus Londinensis ''The Paradisus Londonensis'' (full title ''The Paradisus Londonensis : or Coloured Figures of Plants Cultivated in the Vicinity of the Metropolis'') is a book dated 1805–1808, printed by D.N. Shury, and published by William Hooker.. It consis ...
'' in 1807. In an earlier part of this work, he had used the name ''Leucojum autumnale'' for the plant illustrated in plate 21, but when discussing ''Leucojum pulchellum'', illustrated in plate 74, Salisbury noted the differences between the two species and considered them sufficient to put ''Leucojum autumnale'' into a new genus, ''Acis''. (However, he did not actually use the name ''Acis autumnalis'', which was published in 1829 by Robert Sweet.) Salisbury did not explain the origin of the name ''Acis'' beyond describing it as a "poetic title". It may refer to the myth of Acis and Galatea. Most later botanists did not accept the distinction between ''Leucojum'' and ''Acis''. ''Acis'' was reinstated in 2004, after it was determined on morphological and molecular grounds that the broadly defined genus ''Leucojum'' was
paraphyletic In taxonomy (general), taxonomy, a group is paraphyletic if it consists of the group's most recent common ancestor, last common ancestor and most of its descendants, excluding a few Monophyly, monophyletic subgroups. The group is said to be pa ...
, with ''Acis'' and a more narrowly defined ''Leucojum'' being related as shown the following cladogram. A possible relationship among the species of the genus ''Acis'' is shown in the following cladogram. (In the study, ''Acis ionica'' was treated as "''Acis valentina'' from Greece").


Species

, the
World Checklist of Selected Plant Families The World Checklist of Selected Plant Families (usually abbreviated to WCSP) is an "international collaborative programme that provides the latest peer reviewed and published opinions on the accepted scientific names and synonyms of selected plan ...
accepted nine species, most formerly placed in ''Leucojum''. * '' Acis autumnalis'' (L.) Sweet – W. & W. Central Mediterranean * '' Acis fabrei'' (Quézel & Girerd) Lledó – S.E. France (around Mont Ventoux and River Nesque) * '' Acis longifolia'' J.Gay ex M.Roem. – Corsica * '' Acis nicaeensis'' (Ardoino) Lledó, A.P.Davis & M.B.Crespo – S.E. France (from Nice eastwards) * ''
Acis ionica ''Acis ionica'' is a species of flowering plant in the family Amaryllidaceae, native from south-western Albania to western Greece. It was initially confused with what is now ''Acis valentina'', a species found in Spain near Valencia. Descripti ...
'' Bareka, Kamari & Phitos, syns. ''Leucojum ionicum'' Kit Tan, Mullaj, Sfikas & Strid, ''Acis orientalis'' Strid – SW. Albania to W. Greece * '' Acis rosea'' (F.Martin bis) Sweet – Corsica, Sardinia * ''
Acis tingitana ''Acis tingitana'' is a species of flowering plant in the family Amaryllidaceae, native to north Morocco. Rare in cultivation, it is not fully frost-hardy. Description ''Acis tingitana'' is a bulbous flowering plant. Forms described under the ...
'' (Baker) Lledó, A.P.Davis & M.B.Crespo – Morocco * '' Acis trichophylla'' G.Don – Central & S. Portugal, S. Central & S.W. Spain, Morocco * ''
Acis valentina ''Acis valentina'' is a species of bulbous flowering plant in the family Amaryllidaceae, native to eastern Spain. Its white flowers appear in autumn. It can be grown as an ornamental bulb, but may need protection from hard frost. Description '' ...
'' (Pau) Lledó, A.P.Davis & M.B.Crespo – E. Spain (N. of Valencia) File:Acis autumnalis005.jpg, '' Acis autumnalis'' File:Acis fabrei.jpg , '' Acis fabrei'' File:Acis longifolia in Corsica.jpg, '' Acis longifolia'' File:Acis_nicaeensis.jpg, '' Acis nicaeensis'' File:Acis ionica.jpg , ''
Acis ionica ''Acis ionica'' is a species of flowering plant in the family Amaryllidaceae, native from south-western Albania to western Greece. It was initially confused with what is now ''Acis valentina'', a species found in Spain near Valencia. Descripti ...
'' File:Acis rosea in Corsica.jpg, '' Acis rosea'' File:Leucojum tingitanum 002 GotBot 2016.jpg , ''
Acis tingitana ''Acis tingitana'' is a species of flowering plant in the family Amaryllidaceae, native to north Morocco. Rare in cultivation, it is not fully frost-hardy. Description ''Acis tingitana'' is a bulbous flowering plant. Forms described under the ...
'' File:Leucojum trichophyllum 1.JPG, '' Acis trichophylla'' File:Acis valentina flower.jpg , ''
Acis valentina ''Acis valentina'' is a species of bulbous flowering plant in the family Amaryllidaceae, native to eastern Spain. Its white flowers appear in autumn. It can be grown as an ornamental bulb, but may need protection from hard frost. Description '' ...
''


Cultivation

The species ''Acis autumnalis'' has gained the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (Nort ...
's
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. History The Award of Garden Merit ...
.


References


External links

{{Taxonbar, from=Q2602694 Amaryllidaceae genera